Character designers know a character's basic shapes, silhouette and the overall use of smooth vs sharp shapes help give the character personality, but color scheme is (arguably) the least important element of a character.
Color schemes can be messed with during paintovers as designers see fit without "breaking" the character, and the proper application of shading and highlights can give a flat character a bit more appeal.
Cartoon paintovers are also a fun practice exercise for artists, challenging them to enhance the look of the original characters while keeping their personalities intact underneath the digital paint job.
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