A Grape-Sized Single-Cell Organism

Trace fossils of the Cambrian Gromia sphaerica were recently discovered off the coast of the Bahamas. Although paleontologists had previously assumed that these were multicellular organisms, this particular set of fossils indicate that these giants -- up to 3 cm in diameter -- were single-celled.

Link via Radley Balko

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I've heard that some Slime Molds exist as giant single cell organisms, so these might have acted similarly.

Otherwise, I'm not sure that I've heard of a viable single celled organism that's that large.